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FAMILY CHLOROPIDAE
Paul L.Th. Beuk and Emilia P. Nartshuk Version date: 31-10-2024 10:57

Diagnosis. Chloropidae are small flies, usually 1-5 mm length, rarely till 8 mm. Body stout or elongated, shining black or gray dusted, yellowish red or green with black or red longitudinal stripes. Chaetotaxy of body weakly developed. Head usually with large ocellar triangle; arista sometimes enlarged or thick pubescent. Thorax with vertical ridge on propleuron. Wing with one costal break and a distinct flexure in the long penultimate section of M3+4, opposite or before the middle of cell bm+dm. A few species are wingless or brachypterous.

Biology. Chloropidae occur on meadows, marshes, other low vegetation, sometimes flowers, rarely in forests. They are commonly called grass flies, but only some species have phytophagous larvae, which live in shoots or inflorescences of grasses and sedges, a few are gall formers. Several phytophagous species are known as pests of cereals. Many species are saprophages developing in decaying plants or woods and cones following damage by other insects, or in bird's nest, in fungi, etc. Some species are predators and feed on egg masses of spiders, mantids, and grasshoppers, or on root aphids.

General references. Andersson (1977 [morphology, taxonomy, key to genera]), Chvála et al. (1974 [Lipara]), Duda (1932-1933 [morphology, keys, descriptions]), Ismay (1994 [Neohaplegis, Cryptonevra]), Ismay & Nartshuk (2000 [general, key to genera]), Nartshuk (1966 [morphology], 1972 [biology], 1977 [morphology], 1983 [classification], 1984 [catalogue], 1987 [morphology, classification, evolution, host plants, key to genera for larvae], 1992 [Meromyza], 1994 [zoogeography], 1995 [zoogeography]), Sabrosky (1941 [catalogue of world genera], 1987d [general, key to genera]).

References to the local fauna. Chvála et al. (1974), De Bruyn & von Tschirnhaus (1996), Grijpma & Van der Weerd (1991), De Meijere (1916, 1935, 1939b, 1950a), A.C. Oudemans (1909).

How to quote this page: Beuk, P.L.Th., & E.P. Nartshuk, 2024. Family Chloropidae. In: Beuk, P.L.Th. (Ed.): Checklist of the Diptera of the Netherlands, https://diptera-info.nl/news.php?fam=Chloropidae (date accessed: 27/12/2024).

NL: 147, 1? B: 111 D: 141, 2? UK: 175 World: 2000
 
CHLOROPIDAE
 
CHLOROPINAE
                 
  Camarota Meigen, 1830
  curvipennis (Latreille, 1805)
  = flavitarsis Meigen, 1830
                 
  Cetema Hendel, 1907
  cereris (Fallén, 1820)
  elongatum (Meigen, 1830)
  = nudipes Loew, 1866
  myopinum (Loew, 1866)
  neglectum Tonnoir, 1921 N; informally added by Beuk & Nartshuk (2002: 280
  simile Ismay, 1985 N; informally added by Beuk & Nartshuk (2002: 280
                 
  Chlorops Meigen, 1830
  = Oscinis Latreille, 1804
  subg. Chlorops Meigen, 1830
  adjunctus Becker, 1910 N; informally added by Beuk & Nartshuk (2002: 280
  calceatus Meigen, 1830
  fasciatus Meigen, 1830 Note 1
  frontosus Meigen, 1830
  = scholtzii Egger, 1862
  geminatus Meigen, 1830
  gracilis Meigen, 1830
  hypostigma Meigen, 1830
  laetus Meigen, 1830
  meigenii Loew, 1866 Note 2
  = nasuta (Schrank, 1781): misident.
  = rufescens Oldenberg, 1923
  novakii Strobl, 1902
  obscurellus (Zetterstedt, 1838)
  = brunnipes (Zetterstedt, 1848): misident. sensu auct.
  planifrons (Loew, 1866)
  pumilionis (Bjerkander, 1778)
  = lineatus (Fabricius, 1781): preocc.
  = taeniopus Meigen, 1830
  ringens Loew, 1866
  rossicus Smirnov, 1955 N; informally added by Beuk & Nartshuk (2002: 280
  rufinus (Zetterstedt, 1848)
  scalaris Meigen, 1830
  = didymus (Zetterstedt, 1848)
  serenus Loew, 1866
  speciosus Meigen, 1830
  strigulus (Fabricius, 1794)
  = cingulatus Meigen, 1830
  triangularis Becker, 1910
  troglodytes (Zetterstedt, 1848)
  subg. Sclerophallus Beschovski, 1978
  limbatus Meigen, 1830
  = fulviceps von Roser, 1840
  = brevimanus Loew, 1866
                 
  Cryptonevra Lioy, 1864
  = Haplegis Loew, 1866
  consimilis (Collin, 1932) N; informally added by Beuk & Nartshuk (2002: 280
  diadema (Meigen, 1830)
  flavitarsis (Meigen, 1830)
  = divergens (Loew, 1866)
  nigritarsis (Duda, 1933) N; informally added by Beuk & Nartshuk (2002: 280
                 
  Diplotoxa Loew, 1863
  messoria (Fallén, 1820)
                 
  Epichlorops Becker, 1910
  puncticollis (Zetterstedt, 1848) N; informally added by Beuk & Nartshuk (2002: 280
                 
  Eurina Meigen, 1830
  calva Egger, 1862
  lurida Meigen, 1830
                 
  Eutropha Loew, 1866
  fulvifrons (Haliday, 1833)
  = sulcata (Van der Wulp, 1871)
                 
  Lasiosina Becker, 1910
  albipila (Loew, 1866)
  herpini (Guérin-Méneville, 1843)
  = cinctipes (Meigen, 1830): misident. sensu auct.
                 
  Melanum Becker, 1910
  laterale (Haliday, 1833)
                 
  Meromyza Meigen, 1830
  femorata Macquart, 1835
  mosquensis Fedoseeva, 1960 N; informally added by Beuk & Nartshuk (2002: 281
  nigriseta Fedoseeva, 1960 N; informally added by Beuk & Nartshuk (2002: 281
  nigriventris Macquart, 1835
  ornata (Wiedemann, 1817) N; informally added by Beuk & Nartshuk (2002: 281
  palposa Fedoseeva, 1960 N; informally added by Beuk & Nartshuk (2002: 281
  pratorum Meigen, 1830
  rohdendorfi Fedoseeva, 1974 N; informally added by Beuk & Nartshuk (2002: 281
  saltatrix (Linnaeus, 1758)
  triangulina Fedoseeva, 1960 N; informally added by Beuk & Nartshuk (2002: 281
  variegata Meigen, 1830
  = laeta Meigen, 1838: misident.
                 
  Neohaplegis Beschovski, 1981
  tarsata (Fallén, 1820)
  = laevigata (Fallén, 1820)
                 
  Parectecephala Becker, 1910
  longicornis (Fallén, 1820)
                 
  Platycephala Fallén, 1820
  planifrons (Fabricius, 1798)
  umbraculata (Fabricius, 1794)
                 
  Pseudopachychaeta Strobl, 1902
  approximatonervis (Zetterstedt, 1848)
  oscinina (Fallén, 1823) N; added by Beuk & Lamberts (2023a: 14 Note 3
                 
  Thaumatomyia Zenker, 1833
  = Chloropisca Loew, 1866
  elongatula (Becker, 1910) N; informally added by Beuk & Nartshuk (2002: 280
  glabra (Meigen, 1830)
  hallandica Andersson, 1966
  = obscurella (Zetterstedt, 1838): misident. sensu auct.
  notata (Meigen, 1830)
  = circumdata (Meigen, 1830)
  rufa (Macquart, 1835)
 
OSCINELLINAE
                 
  Aphanotrigonum Duda, 1932
  fasciella (Zetterstedt, 1855) N; informally added by Beuk & Nartshuk (2002: 282
  femorellum Collin, 1946 N; informally added by Beuk & Nartshuk (2002: 282
  inerme Collin, 1946 N; informally added by Beuk & Nartshuk (2002: 282
  meijerei (Duda, 1933)
  nigripes (Zetterstedt, 1848)
  = brachypterum (Zetterstedt, 1848)
  trilineatum (Meigen, 1830) Note 4
  = annuliferum (Zetterstedt, 1848)
  = beschovskii Dely-Draskovits, 1981
                 
  Calamoncosis Enderlein, 1911
  subg. Calamoncosis Enderlein, 1911
  aprica (Meigen, 1830)
  aspystylina Duda, 1935 N; informally added by Beuk & Nartshuk (2002: 282
  duinensis (Strobl, 1909)
  minima (Strobl, 1893)
  = nigricornis (von Roser, ): nomen dubium
  subg. Rhaphiopyga Nartshuk, 1971
  glyceriae Nartshuk, 1958 N; informally added by Beuk & Nartshuk (2002: 282
                 
  Conioscinella Duda, 1929
  frontella (Fallén, 1820)
  gallarum (Duda, 1933) N; informally added by Beuk & Nartshuk (2002: 282
  mimula Collin, 1946 N; informally added by Beuk & Nartshuk (2002: 282
  sordidella (Zetterstedt, 1848)
  = flavifrons Duda, 1933: sensu De Meijere, 1935
  zetterstedti Andersson, 1966 N; informally added by Beuk & Nartshuk (2002: 282
                 
  Dicraeus Loew, 1873
  subg. Dicraeus Loew, 1873
  ingratus (Loew, 1866)
  raptus (Haliday, 1838) N; informally added by Beuk & Nartshuk (2002: 282
  tibialis (Macquart, 1835)
  = pallidiventris (Macquart, 1835)
  = bipunctata (Macquart, 1835)
  subg. Oedesiella Becker, 1910
  fennicus Duda, 1933 N; informally added by Beuk & Nartshuk (2002: 282
  subg. Paroedesiella Anonymous, 1937
  styriacus (Strobl, 1898)
  vagans (Meigen, 1838)
  = xanthopygus (Strobl, 1909)
                 
  Elachiptera Macquart, 1835
  bimaculata (Loew, 1845)
  brevipennis (Meigen, 1830)
  cornuta (Fallén, 1820)
  diastema Collin, 1946 N; added by De Bruyn & von Tschirnhaus (1996: 43, 195
  megaspis (Loew, 1858) N; informally added by Beuk & Nartshuk (2002: 282
  rufifrons Duda, 1932
  scrobiculata (Strobl, 1901) N; informally added by Beuk & Nartshuk (2002: 282
  tuberculifera (Corti, 1909) N; informally added by Beuk & Nartshuk (2002: 282
                 
  Eribolus Becker, 1910
  gracilior (De Meijere, 1918)
  hungaricus Becker, 1910
  = plana (De Meijere, 1918)
  nana (Zetterstedt, 1838)
  = sudeticus Becker, 1910
  = leptomyzoides (De Meijere, 1918)
  slesvicensis Becker, 1910
                 
  Gampsocera Schiner, 1862
  numerata (Heeger, 1858) N; added by Dek et al. (2016: 11 Note 5
                 
  Gaurax Loew, 1863
  dubius (Macquart, 1835)
  fascipes Becker, 1910 listed by Nartshuk (2020: 362
  flavomaculatus (Duda, 1933) listed by Nartshuk (2020: 362
  maculipennis (Zetterstedt, 1848) N; informally added by Beuk & Nartshuk (2002: 283
  niger Czerny, 1906 N; informally added by Beuk & Nartshuk (2002: 283
                 
  Hapleginella Duda, 1933
  laevifrons (Loew, 1858)
                 
  Lasiambia Anonymous, 1937
  = Fiebrigella Duda, 1921
  = Lasiamba Sabrsoky, 1941: incorrect spelling by Beuk & Nartshuk, 2002
  baliola (Collin, 1946) N; informally added by Beuk & Nartshuk (2002: 283
  brevibucca (Duda, 1932) N; informally added by Beuk & Nartshuk (2002: 283
  palposa (Fallén, 1820)
                 
  Lasiochaeta Corti, 1909
  = Melanochaeta Bezzi, 1906
  pubescens (Thalhammer, 1898) N; added as species of Elachiptera by De Bruyn & von Tschirnhaus (1996: 43, 195
                 
  Lipara Meigen, 1830
  lucens Meigen, 1830
  pullitarsis Doskočil & Chvála, 1971
  rufitarsis Loew, 1858
  = tomentosa (Macquart, 1835): sensu Becker
                 
  Microcercis Beschovski, 1978
  = Incertella Sabrosky, 1980 Note 6
  albipalpis (Meigen, 1830)
  antennata (Collin, 1946) listed by Nartshuk (2020: 362
  kerteszi (Becker, 1910)
  trigonella (Duda, 1933) N; informally added by Beuk & Nartshuk (2002: 283
  zuercheri (Duda, 1933) N; informally added by Beuk & Nartshuk (2002: 283), formally listed by Nartshuk (2003: 32
                 
  Oscinella Becker, 1909
  angularis Collin, 1946 N; informally added by Beuk & Nartshuk (2002: 283
  angustipennis Duda, 1933 [T] 
  capreolus (Haliday, 1838) Note 7
  = aterrima (Strobl, 1880)
  frit (Linnaeus, 1758)
  = fumipennis (Meigen, 1830)
  = nigrita (Meigen, 1838)
  maura (Fallén, 1820)
  = albiseta (Meigen, 1830)
  nigerrima (Macquart, 1835) listed by Nartshuk (2020: 362
  nitidissima (Meigen, 1838) N; informally added by Beuk & Nartshuk (2002: 283
  pusilla (Meigen, 1830) Note 8
  trochanterata Collin, 1946 N; informally added by Beuk & Nartshuk (2002: 283
                 
  Oscinimorpha Lioy, 1864
  albisetosa (Duda, 1932)
  = hollandica (Duda, 1932)
  arcuata (Duda, 1932) N; informally added by Beuk & Nartshuk (2002: 284
  longirostris (Loew, 1858) N; material in ZMAN examined by Nartshuk  Note 9
  minutissima (Strobl, 1900)
  = longirostris (Loew, 1858): misident. sensu De Meijere, 1916
  sordidissima (Strobl, 1893)
                 
  Oscinisoma Lioy, 1864
  = Oscinosoma Enderlein, 1911:
  cognatum (Meigen, 1830)
                 
  Polyodaspis Duda, 1933
  = Polydaspis Duda, 1933: incorrect spelling by Beuk & Natshuk (2002)
  ruficornis (Macquart, 1835)
                 
  Rhodesiella Adams, 1905
  plumiger (Meigen, 1830)
                 
  Rhopalopterum Duda, 1929
  anthracinum (Meigen, 1830)
  = fasciola (Meigen, 1830): misident.
  atricillum (Zetterstedt, 1838)
  = nitidissima (Meigen, 1838): misident.
  atricorne (Zetterstedt, 1838) N; informally added by Beuk & Nartshuk (2002: 284
  brunneipenne Beschovski & Lansbury, 1987 N; informally added by Beuk & Nartshuk (2002: 284
  fasciola (Meigen, 1830) N; informally added by Beuk & Nartshuk (2002: 284
  femorale (Collin, 1946) N; informally added by Beuk & Nartshuk (2002: 284
                 
  Siphonella Macquart, 1835
  oscinina (Fallén, 1820)
                 
  Siphunculina Rondani, 1856
  aenea (Macquart, 1835)
                 
  Speccafrons Sabrosky, 1980
  halophila (Duda, 1933)
                 
  Trachysiphonella Enderlein, 1936
  ruficeps (Macquart, 1835)
  = pygmaea (Meigen, 1838) Note 10
  = flavella (Zetterstedt, 1848)
  scutellata (von Roser, 1840)
  = pumilionis (Bjerkander, 1778): misident.
  = pumilio (Zetterstedt, 1838): misident. sensu auct.
                 
  Tricimba Lioy, 1864
  subg. Nartshukiella Beschovski, 1981
  albiseta Dely-Draskovits, 1983 N; informally added by Beuk & Nartshuk (2002: 284
  cincta (Meigen, 1830) Note 11
  = sulcella (Zetterstedt, 1848): misident.?
  subg. Tricimba Lioy, 1864
  lineella (Fallén, 1820)
 
Doubtful species
 
  Thaumatomyia Zenker, 1833    
  trifasciata (Zetterstedt, 1848) added by De Meijere (1950a: 12 Note 12
 
Deleted species
 
  Lipara Meigen, 1830  
  similis Schiner, 1854    
 
Notes
Note 1 De Meijere (1939b: 167) listed C. dasycerus Loew, 1866, and C. horridus Becker, 1910, as synonyms of C. fasciatus but these are considered as valid species by Dely-Draskovits (1978). However, these species have not yet been found in the Netherlands.
Note 2 The name Chlorops meigenii is preoccupied by Chlorops meigenii Fallén, 1823 (now in Agromyzidae) but the International Commission on Zoological Nomenclature ruled that the current usage of this name in the Chloropidae was to be conserved (ICZN, 2003: 166).
Note 3 Material examined: NOORD-HOLLAND: Castricum (Noordhollands Duinreservaat, Zanderij [GPS: 52.548N, 4.651E], 2 [f]: 11.viii.2022, leg. S. Lamberts [NHMM ALC]).
Note 4 Again added as beschovskii by De Bruyn & von Tschirnhaus (1996: 43, 195).
Note 5 Material examined: LIMBURG: Cadier en Keer (oostzijde Riesenberg, 1 [m]: 25.iii.2013, leg. P.L.Th. Beuk [NHMM ALC]), Maastricht (Natuurhistorisch Museum Maastricht, indoors, 1 [f]: 11.vi.2013, leg. P.L.Th. Beuk [NHMM ALC]).
Note 6 Synonymy pointed out by Foster (2024: 405).
Note 7 Transferred from Melanochaeta by Nartshuk & Andersson (2013: 126).
Note 8 This species was last recorded by A.C. Oudemans (1909: 22) but it was not listed by De Meijere (1939b), possibly because for some time it was considered to be as synonym of O. frit. Recent material was examined by Nartshuk so this species is maintained on the Dutch list.
Note 9 The material recorded as O. longirostris by De Meijere (1916: 310) belongs to O. minutissima but additional material belonging to O. longirostris is present in ZMAN.
Note 10 Synonymy with ruficeps established by Nartshuk & Andersson (2013: 149).
Note 11 T. sulcella was traditionally placed as synonym of T. cincta but re-instated as valid species by Dely-Draskovits (1983: 346). De Meijere (1939b: 167) lists T. cincta but does not give T. sulcella as synonym, although that species was recorded earlier, e.g., in De Meijere (1916: 311). Until material has been checked T. sulcella will be listed as misidentification of T. cincta.
Note 12 De Meijere (1950a: 12) recorded reared material from Cichorium. However, this is a species of very humid places, so the record is considered doubtful. Possibly the material belongs to T. hallandica.
 
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Incertella was the last (sub)genus to be edited on 31-10-2024 10:57
Microcercis zuercheri was the last (sub)species to be edited on 31-10-2024 10:56















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